Aboard a submarine, Julien, Pancho and Maurice consider their next move; while being held captive, Mort plans to take down the tyrannical new king.
Clover hopes Sage's guru can assist with a plot to oust Koto, who has made Mort his jester.
Mort leads an effort to get rid out Koto; Julien, Pancho and Maurice are faced with a monster; Sage's guru isn't what Clover was expecting.
As Julien and his pals travel back to Madagascar, their submarine has some trouble; Mort puts on a play to disguise an escape.
Julien and the guys work with sharks in hopes of forming an army; a map leads Sage and Clover into danger; Mort's plan hits a snag.
Disgruntled Russian space chimps could be Julien's best hope for battling Koto; Mort finds Timo locked in his pod.
Clover reconsiders her decision to join Pam's cult; Mort tries to contact Julien and company, who are now in trouble with the chimps.
While in disguise, Julien and friends try to catch a ride on a Foosa ship; Clover is presented with a baffling new challenge.
Maurice and Julien are faced with a harrowing battle at gladiator school; Mort goes to another dimension; Sage convinces Clover that there is still good in Koto.
Bell-shaped creatures deem Maurice their chosen one; Julien is stuck in limbo and must stand trial; Mort fights Morticus Khan.
Karl and Julien try to stop Koto's wedding to the Crocodile Princess; Mort fights to bring "Smart Mort" back.
Koto wants to kick off the wedding with a sacrifice; Sage and Clover unleash their weapon; Maurice, Mort and Pancho show up with reinforcements.
Julien turns to Mort's army, the bell snails, the space chimps, Clover and other loyal friends for help.